The country of Wanokuni is enclosed by towering walls and houses a colossal tree at its core. Within its borders, two siblings named Mana and Shino coexist in harmony alongside their acquaintances and locals. Unfortunately, this tranquility is swiftly interrupted when two unfamiliar women abruptly appear from beyond Wanokuni's borders. Chaos ensues as Wanokuni is under siege! As Mana helplessly witnesses his comrades perish, he discovers that Wanokuni is, in fact, an enormous living entity, and the invaders have arrived on a rival beast. This marks the beginning of the siblings' journey, as they engage in a momentous conflict that will determine the future of their homeland!